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The biodiversity increases when one moves from the pioneer to the climax stage. What could be the explanation?

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The biodiversity increases when one moves from the pioneer to the climax stage. What could be the explanation?

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In the successive serial stages there is a change in the diversity of species of organisms, increase in the number of species and organisms as well as an increase in the total biomass. Description of ecological succession usually focuses on changes in vegetation. However, these vegetational changes in turn affect food and shelter for various types of animals. Thus, as succession proceeds, the numbers and types of animals and decomposers also change.

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