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Select one of the following pairs of important features distinguishing Gnetum from Cycas and Pinus and showing affinities with angiosperms

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Select one of the following pairs of important features distinguishing Gnetum from Cycas and Pinus and showing affinities with angiosperms

(1) perianth and two integuments

(2) absence of resin duct and leaf venation

(3) embryo development and apical meristem

(4) presence of vessel elements and absence of archegonia

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(4) presence of vessel elements and absence of archegonia

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