All of the following statements concerning the Actinomycetes filamentous soil bacterium Frankia are correct except that Frankia:

(1) Forms specialized vesicles in which the nitrogenase is protected from oxygen by a chemical barrier involving triterpene hopanoids

(2) Can induce root nodules on many plant species

(3) Like Rhizobium, it usually infects its host plant through root hair deformation and stimulates cell proliferation in the host’s cortex

(4) Cannot fix nitrogen in the free-living state